[Chen, Yanping;Evans, Jay D.;Zhou, Liang;Boncristiani, Humberto;Pettis, Jeffery S.] ARS, USDA, Bee Res Lab, Beltsville, MD 20705 USA.^[Kimura, Kiyoshi] Natl Inst Livestock & Grassland Sci, Res Team Anim Breeding, Honeybee Res Grp, Tsukuba, Ibaraki, Japan.^[Xiao, Tieguang] Hunan Agr Univ, Coll Plant Protect, Dept Entomol, Changsha, Hunan, Peoples R China.^[Litkowski, A. M.] Univ Maryland, Coll Life Sci, Dept Biol, College Pk, MD 20742 USA.