We thank the journal reviewers for helpful comments and an anonymous reviewer for English improvements. The current study was supported by the Chinese Academy of Sciences (Hundred Talent Program Y62302, Institute Talent Program Y45Z04, and Key Strategic Program ZDRW-ZS-2017-3-2), the State Key Laboratory of Freshwater Ecology and Biotechnology (Grant 2016FBZ10) and the Hunan Provincial Innovation Foundation for Postgraduate (CX2017B349).