Conceptualization, data curation, and supervision, G.H., R.L. and X.H.; investigation and methodology, G.H., L.W., R.L. and F.C.; resources and software, R.L. and J.Y.; writing and editing, G.H. and R.L.; funding acquisition, Y.Y., R.L. and X.H. All authors have read and agreed to the published version of the manuscript. This work was jointly supported by the National Key Research and Development Program (2021YFD1300201, 2021YFD1301004), the National Natural Science Foundation of China (U20A2055, 32172761), the Natural Science Foundation of Hunan Province (2022JJ40532), the Key Science and Technology Special Program of Yunnan Province (202202AE090032), and the Open Fund of Key Laboratory of Agro-ecological Processes in Subtropical Region, Chinese Academy of Sciences (ISA2021103, ISA2023201).