This work was financially supported by the Research of The Major Science and Technology Special Project in Hunan, China (2017NK1030) and the National Natural Science Foundation of China (31572626). This research complied with ethical standards. The entire experiment was conducted according to the guidelines established by the National Institutes of Health. To minimize suffering, all experimental fish were anesthetized with eugenol (1:12,000; Shanghai Reagent Corporation, Shanghai, China) before sampling. There is no conflict of interest declared in this article.