Conceptualization, X.X. and Y.Y.; methodology, Y.S., R.L., L.Z., H.L., C.P. (Can Peng) and M.Q.; software, Y.S.; validation, X.X. and Y.S.; formal analysis, Y.S.; investigation, X.X. and Y.S.; resources, X.X.; data curation, Y.S.; writing—original draft preparation, Y.S.; writing—review and editing, X.X., Q.P., H.L., and L.Z.; visualization, Y.S.; supervision, X.X.; project administration, Y.S., C.P. (Changfeng Peng), C.P. (Can Peng), S.W., Q.P. and Y.W.; funding acquisition, Y.Y., X.X., R.L. and L.Z. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Natural Science Foundation of China (32130099), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDA28110100), the Natural Science Foundation of Hunan Province (2022JJ40532), the Open Fund of Key Laboratory of Agroecological Processes in Subtropical Region, Chinese Academy of Sciences (ISA2021103), the Hunan Provincial Natural Science Foundation of China (2021JJ30171), and the Research Foundation of Education Bureau of Hunan Province, China (21A0584).
