Conceptualization, H.Z., H.X. and Y.Z.; methodology, H.Z.; validation, H.Z. and X.T.; investigation, H.Z. and J.W.; writing—original draft preparation, H.Z.; writing—review and editing, H.Z. and H.X.; supervision, Y.Z.; project administration, H.X. and Q.T.; funding acquisition, Q.T. All authors have read and agreed to the published version of the manuscript. This research was fund by the Earmarked Fund for China Agriculture Research System (Grand No. CARS-01-27).