Conceptualization, T.Z.; methodology, M.Z. and Y.B.; software, Y.S.; validation, M.Z., Y.S. and J.A.; formal analysis, Y.S.; investigation, T.Z.; resources, T.Z.; data curation, T.Z.; writing—original draft preparation, M.Z. and Y.B.; writing—review and editing, T.Z. and Q.C.; visualization, Y.S.; supervision, T.Z.; project administration, T.Z.; funding acquisition, T.Z. All authors have read and agreed to the published version of the manuscript. This research was funded by National Key R&D Program of China (2021YFD1301004), the Central Public-interest Scientific Institution Basal Research Fund (Y2017CG36), National Natural Science Foundation of China (31470122) and Special fund for science and technology innovation project of Chinese Academy of Agricultural Sciences (ASTIP-IAS08).