This research was funded by National Natural Science Foundation of China (No. 31672457), Hunan Provincial Science and Technology Department (2020NK2004, 2021JJ30008) and Double first-class construction project of Hunan Agricultural University (SYL201802003, YB2018007, CX20190497, CX20190524). CRediT authorship contribution statement Liu Zhuangzhuang: Conceptualization, Methodology, Software, Data curation, Writing – original draft, Writing – review & editing. Fang Jun: Conceptualization, Writing – original draft, Supervision, Project administration, acquisition.