本研究由国家重点研发计划项目(2017YFD0200703), 国家自然科学基金区域创新发展联合基金项目(U19A2050), 湖南省自然科学基 金青年项目(2019JJ50233, 2019JJ50337)和湖南省教育厅重点项目(20A250)资助。 This study was supported by the National Key Research and Development Project of China (2017YFD0200703), the Joint Funds of the National Natural Science Foundation of China (U19A2050), the Hunan Natural Science Foundation Project (2019JJ50233, 2019JJ50337), and the Key Project of Hunan Provincial Education Department (20A250). * 通信作者(Corresponding author): 王英姿, E-mail: 83075688@qq.com 第一作者联系方式: E-mail: chtian12@126.com Received (收稿日期): 2020-05-24; Accepted (接受日期): 2020-09-13; Published online (网络出版日期): 2020-10-05. URL: https://kns.cnki.net/kcms/detail/11.1809.S.20200930.1634.008.html