We thank Prof. Chun-Yun Guan (Hunan Agricultural University, Changsha) for providing rapeseed materials. We thank Shanghai Applied Protein Technology (Shanghai, China), Biotree (Shanghai, China), and Novogene Biotechnology Company (Beijing, China) for valuable technical help. This study was supported by the National Natural Science Foundation of China (U21A20236, 32072664), Natural Science Foundation of Hunan Province (2021JJ0004), China Agriculture Research System, and the Hunan Postgraduate Scientific Research Innovation Project (CX20190505).