1 Hunan Agriculture University / Upland Crop Research Institute, Hunan Agriculture University, Changsha 410128, Hunan, China; 2 College of Biology and Environment Engineering, Binzhou University, Binzhou 256600, Shandong, China; 3 Shandong Peanut Research Institute / National Peanut Engineering Technology Research Center, Qingdao 266100, Shandong, China; 4 Linyi Academy of Agricultural Sciences, Linyi 276000, Shandong, China; 5 Shandong Academy of Agricultural Sciences / Key Laboratory of Crop Genetic Improvement and Ecological Physiology of Shandong Province, Jinan 250100, Shandong, China 本研究由国家重点研发计划项目(2018YFD1000906), 财政部和农业农村部国家现代农业产业技术体系建设专项(CARS-13), 山东省重 大科技创新工程项目(2019JZZY010702)和山东省农业科学院农业科技创新工程项目(CXGC2022A10)资助。 This study was supported by the National Key Research and Development Program of China (2018YFD1000906), the China Agriculture Research System of MOF and MARA (CARS-13), the Major Scientific and Technological Innovation Projects in Shandong Province (2019JZZY010702), and the Agricultural Science and Technology Innovation Project of Shandong Academy of Agricultural Sciences (CXGC2022A10). * 通信作者(Corresponding authors): 万书波, E-mail: wanshubo2016@163.com; 李林, E-mail: Lilindw@163.com ** 同等贡献(Contributed equally to this work) 第一作者联系方式: 刘俊华, E-mail: liujh516@163.com Received (收稿日期): 2022-02-21; Accepted (接受日期): 2022-06-07; Published online (网络出版日期): 2022-07-09. URL: https://kns.cnki.net/kcms/detail/11.1809.S.20220707.1530.018.html This is an open access article under the CC BY-NC-ND license (http://creativecommons.org/licenses/by-nc-nd/4.0/).