Writing—original draft preparation, X.F.; writing—review and editing, X.F. and S.D.; data curation, X.F.; methodology, X.C. and Z.D.; software, H.X., H.K. and F.C.; supervision, X.C., R.W. and S.D.; project administration, Y.S. and S.D.; funding acquisition, Y.S. and S.D. All authors have read and agreed to the published version of the manuscript. This study was supported by the National Natural Science Foundation of China (31501543, 32272257), Training Program for Excellent Young Innovators of Changsha (KQ1905025), Natural Science Foundation of Hunan Province, China (2022JJ30330), and the Agricultural Science and Technology Innovation Project of Hunan Province, China (2021CX05, 2022CX44).