Conceptualization, Y.X. and J.F.; methodology, S.L.; software, Y.M.; validation, S.L., Y.X. and H.L.; formal analysis, S.L.; investigation, R.C.; resources, Z.C.; data curation, B.Y. and S.L.; writing—original draft preparation, S.L.; writing—review and editing, S.L.; visualization, S.L.; supervision, Y.L.; project administration, Y.X. and J.F.; funding acquisition, Y.X. and J.F. All authors have read and agreed to the published version of the manuscript. The study was supported by the National Natural Science Foundation of China (No. 41807135, No. 41877491 and No. 31800076), the Natural Science Foundation of Hunan province, China (No. 2019JJ50220), the Scientific Research Fund of Hunan Provincial Education Department (No. 18B107), Hunan Provincial Science and Technology Department (2018WK4025, 2020NK2004, 2020ZL2004), and Double first-class construction project of Hunan Agricultural University(SYL201802003, YB2018007, CX20190497).
