This work was supported by the Hunan Provincial Key Laboratory for Synthetic Biology of Traditional Chinese Medicine (No: HCSW2023-01), the Key Scientific Research Fund of Hunan Provincial Education Department (No: 21A0138), the Project of Hunan Natural Science Foundation (No: 2022JJ30305), the Horticulture’s Open program of Hunan Agricultural University (No: 2021YYXK002), Postgraduate Scientific Research Innovation Project of Hunan Province (No: CX20220689) and Postgraduate Scientific Research Innovation Project of Hunan Agriculture University (No: 2022XC062).