Conceptualization, J.Z.; methodology, X.S.; software, L.H.; validation, Z.Z. (Zhuo Zhang) and Z.Z. (Zhanghong Zhang); formal analysis, X.S.; investigation, Y.L.; resources, D.Z.; data curation, J.Z.; writing—original draft preparation, J.Z.; writing—review and editing, X.S.; visualization, Z.Z. (Zhuo Zhang); supervision, Y.Z.; project administration, X.S.; funding acquisition, X.S. All authors have read and agreed to the published version of the manuscript. This work was supported by the National Natural Science Foundation of China (Nos. 32030088, 32272535, 32072383, 31901854) and the Agriculture Research System of China (No. CARS-23-D-02).