This research was supported by the National Research and Development Program ( 2017YFD0200502 ), the National Natural Science Foundation of China ( 31420103919 and 31672032 ), the Beijing Nova Program ( Z171100001117039 ), the Science and Technology Innovation Program of the Chinese Academy of Agricultural Sciences ( AAS-ASTIP-IVFCAAS ), and the Beijing Key Laboratory for Pest Control and Sustainable Cultivation of Vegetables .