This research was supported by the Natural Science Foundation of China (No. 31272248) and the project of graduate innovation in Hunan province (No. CX2017B364). The authors thank Zhenzhen Zhang of the Foreign Language College of Hunan Agricultural University and Professor Ninghu Su of the College of Science and Engineering, The Centre for Tropical Water & Aquatic Ecosystem Research (TropWATER) and the Cairns Institute, James Cook University for the revision of the English writing. We are grateful the guidance of the editor, Mahmood Farooq, and other reviewers whose constructive criticism greatly helped us to improve the quality of this manuscript.
