H.Y.: Formal analysis, Methodology, Investigation, writing—original draft preparation, Validation, Visualization. Q.T.: Conceptualization, Validation—review and editing. J.Z. (Jun Zhou): Investigation, Writing—review and editing. J.Z. (Jing Zhang): funding acquisition, Writing—review and editing. X.H.: Investigation, Validation. S.P.: Formal analysis, Validation. H.L.: Formal analysis. Z.D.: Visualization. X.L.: Investigation. Y.T.: Formal analysis. G.G.: Formal analysis. Y.H.: Conceptualization, Validation, Writing—review and editing, Visualization, Supervision, Funding acquisition, Resources. X.Z.: Conceptualization, Validation, Formal analysis, Funding acquisition, Investigation. All authors have read and agreed to the published version of the manuscript. This research was funded by the China Postdoctoral Science Foundation (2023M741143), Hunan Province Natural Science Foundation (2021JJ30320), the Scientific Research Fund of Hunan Provincial Education Department (23B0228), the Research Innovation Project of Postgraduates in Hunan Agricultural University (CX20230706), and the Seed Industry Innovation Project of Hunan province of China (2021NK1005). Postgraduate Scientific Research Innovation Project of Hunan Province (QL 20220172).