Conceptualization, Z.L., L.B. and L.P.; methodology, Z.L., H.L., J.W. and W.C.; formal analysis, Z.L., H.L. and L.P.; investigation, Z.L., J.W. and L.P.; resources, L.B. and L.P.; writing—original draft preparation, Z.L., L.B. and L.P.; writing—review and editing, L.B. and L.P.; supervision, L.B. and L.P.; funding acquisition, L.B. and L.P. All authors have read and agreed to the published version of the manuscript. This research was financially supported by the National Natural Science Foundation of China (32130091, 31901905), Young Elite Scientists Sponsorship Program by CAST (2021QNRC001), the Natural Science Foundation of Hunan Province, China (2021JJ20002), the Science and Technology Innovation Program of Hunan Province (2021RC3087), and China Agriculture Research System (CARS-16-E19).