The study was financially supported by the National Natural Science Foundation of China (Grant No.51909089), Natural Science Foundation of Hunan Province, China (Grant No.2020JJ5252, 2021JJ40091), China Postdoctoral Science Foundation (Grant No. 2019M662781), Science Foundation for Young Scholars of Hunan Agricultural University (19QN35), and Hunan Provincial Innovation Foundation for Postgraduate (CX20200663).