The authors thank Dr Min Wang (South China Agricultural University, China), Mr Ming-Wei Liao (Hunan Agricultural University, China), Mr Wen-Wei Zhang, Mr Min Wang and Mr Ming-Jian Wang (Hunan Wuyunjie National Nature Reserve Administration) for their help on collection in the field. The authors also thank Yume Imada (Ehime University, Japan) for her careful reading of our manuscript and for many constructive comments. This study was supported by the Biodiversity Survey and Assessment Project of the Ministry of Ecology and Environment, China (2019HJ2096001006), JSPS and NSFC under the Japan-China Scientific Cooperation Program, and JSPS KAKENHI Grant Number JP16H05766, and Double first-class construction project of Hunan Agricultural University.