Conceptualization, H.J. and H.L.; Software, H.J. and H.L.; Investigation, H.L.; Writing—original draft, H.J.; Writing—review & editing, H.L.; Supervision, H.L.; Funding acquisition, H.J. and H.L. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Social Science Foundation Project of China (Grant No. 24BJY165), Postgraduate Scientific Research Innovation Project of Hunan Province (Grant No. CX20240678) and the Postgraduate Scientific Research Innovation Project of Hunan Agricultural University (Grant No. 2024XKC072).