National Key R & D Program of China, China [2019YFC1604903, 2019YFC1604905]; National Natural Science Foundation of China, China [21808052]; Hunan Provincial Natural Science Foundation, China [2022JJ50022]; Science and Technology Innovation Program of Hunan Province, China [2022RC1170]; Training Program for Excellent Young Innovators of Changsha, China [kq2206037]; Double First -Class Construction Project of Hunan Agricultural University, China [SYL201802002]