Conceptualization, H.X., X.Y. and R.W.; methodology, H.X., X.Y. and R.W.; software, X.Y., S.L., J.C., Q.L. and F.X.; validation, H.X., X.Y. and Q.L.; formal analysis, H.X., X.Y., S.L., J.C., Q.L., L.S., W.S., F.X., L.D. and R.W.; investigation, X.Y. and Q.L.; resources, H.X., L.D. and R.W.; data curation, H.X. and R.W.; writing—original draft preparation, H.X. and X.Y.; writing—review and editing, H.X., X.Y., S.L., J.C., Q.L., L.S., W.S., F.X., L.D. and R.W.; visualization, H.X. and X.Y.; supervision, R.W. and L.D.; project administration, R.W. and L.D.; funding acquisition, H.X. All authors have read and agreed to the published version of the manuscript. This research was funded by the Joint Fund for Scientific and Technological Research and Development of Henan (222301420110) and the Distinguished Young Scholars from the Henan Academy of Agricultural Sciences (2022JQ04).