Conceptualization, J.J. and S.C.; methodology, S.C. and P.W.; software, J.J. and P.W.; validation, P.W., J.J. and W.C.; formal analysis, H.L.; investigation, L.Y.; resources, Z.F. and Y.Z.; data curation, W.C. and J.J.; writing—original draft preparation, P.W. and W.C.; writing—review and editing, J.J. and S.C.; visualization, J.J. and W.C.; supervision, H.L.; project administration, M.Z. and Y.W.; funding acquisition, J.J. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Key Research and Development Program (2022YFD1200502), National Natural Science Foundation of China (32002034), the Science and Technology Innovation Program of the Chinese Academy of Agricultural Sciences (CAAS-ASTIP-2013-IVFCAAS), and the Modern Agro-Industry Technology Research System (CARS-25-B-01), and State Key Laboratory of Vegetable Biobreeding, Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Beijing 100081, China.