This work was supported by the National Natural Science Foundation of China (Nos. 31472017, 31272339), Project of Science and Technology Department of Hunan Province (No.2014FJ2003), Tropical Disease Control and Research, Ministry of Education in China (2018kfkt03), Natural Science Foundation of Hunan Province (2020JJ4427), and Key Project of Education Department of Hunan Province (18A024).