Conceptualization, M.X.; data curation, M.X., Q.D., S.L. (Si Liu), H.W., J.X., J.Z., X.Y. and S.L. (Shaoming Li); formal analysis, M.X., P.W., S.L. (Si Liu), H.W., J.X. and X.Y.; funding acquisition, R.S. and Z.X.; investigation, M.X. and J.Z.; methodology, M.X., P.W., and R.S.; resources, J.X., R.S. and Z.X.; software, H.W. and J.Z.; supervision, S.L. (Shaoming Li) and Z.X.; validation, Z.Z.; visualization, M.X.; writing—original draft, M.X.; writing—review and editing, Z.Z. and R.S. All authors have read and agreed to the published version of the manuscript. This research was funded by the Earmarked Fund for China Agriculture Research System (grant number: CARS-45), the Hunan Province Science and Technology Plan project (grant number: 2016NK2177), and the Changsha Natural Science Foundation for Changsha Science and Technology Bureau (grant number: kq2202354).