This work was financially supported by the National Key Research and Development Program of China (2021YFD1700200), the National Natural Science Foundation of China (32202607, 32302676), the Hunan Provincial Natural Science Foundation (2023JJ40392), the Hunan Provincial Fund for Agricultural Science and Technology Innovation (2023CX47, 2023CX45), and the Special Fund for the Construction of Modern Agricultural Industry Technology System (CARS-22-G-11). CRediT authorship contribution statement Changyu Fang: Writing – original draft, Visualization, Methodology, Formal analysis, Data curation. Peng Li: Writing – review & editing, Methodology. Jianglin Zhang: Methodology. Yanhong Lu: Project administration, Investigation. Youyun Tang: Project administration, Investigation. Naimei Tu: Writing – review & editing, Supervision, acquisition. Yulin Liao: Project administration, Investigation, acquisition, Writing – review & editing. Jun Nie: Resources, Project administration,