This study was financially supported in part, by the National Key Research & Development Program of China (2017YFD0200100, 2017YFD0200103); National Natural Science Foundation of China (Grant No.31101596, 31372130); Hunan Provincial Recruitment Program of Foreign Experts; and the National Oilseed Rape Production Technology System of China; '2011 Plan' supported by The Ministry of Education of China