This work was supported by the research fund of Zhejiang Science and Technology Major Program on Agricultural New Variety Breeding (2021C02064-6). CRediT authorship contribution statement Biao Luo: Writing – original draft, Visualization, Validation, Investigation, Data curation. Xianwen Zhang: Validation, Resources, Investigation, acquisition, Data curation. Fang Wang: Visualization, Validation. Yan Wang: Methodology, Formal analysis. Wei Wu: Resources. Chaoyang Lin: Methodology, Formal analysis. Liqun Rao: Writing – review & editing, Supervision, Project administration, Conceptualization. Qiming Wang: Writing – review & editing, Supervision, Conceptualization.