This work was supported by National Natural Science Foundation of China ( 31601525 ), Natural Science Foundation of Changsha ( kq2208075 ), Natural Science Foundation of Hunan Province ( 2019JJ50256 ), National Natural Science Foundation of China ( 32272257 ), The Agricultural Science and Technology Innovation Project of Hunan Province ( 2022CX44 ), Funds for Distinguished Young Scientists of Changsha ( KQ1905025 ).