This work was supported by the National Natural Science Foundation of China ( 32372237 , 31801319 ), the National Key Research and Development Program of China ( 2024YFD2300301 ), the Science and Technology Innovation Program of Hunan Province ( 2023RC3153 ), the Natural Science Foundation for Distinguished Young Scholars of Changsha City ( kq2209016 ), and the Key Scientific Research Project of the Hunan Provincial Department of Education ( 24A0160 ). CRediT authorship contribution statement Cheng Huang: Writing – review & editing, Visualization, Software, acquisition. Pan Long: Resources. Qiwen Hou: Investigation, Data curation. Ying Xu: Writing – review & editing, Writing – original draft, Visualization, Supervision, Software, Methodology, acquisition, Formal analysis, Data curation, Conceptualization. Mutian Gao: Investigation. Jinbiao Xiang: Investigation, Data curation. Sheng Chen: Investigation. Pufan Shao: Investigation. Zhiqiang Fu: Resources.