This work was supported by the China Agriculture Research System of MOF and MARA (CARS-19), the Science and Technology Innovation Project of the Chinese Academy of Agricultural Sciences (CAAS-ASTIP-TRICAAS), the National Natural Science Foundation of China (Grant No.:32202546), the Central Public-interest Scientific Institution Basal Research Fund for Chinese Academy of Agricultural Sciences (Grant No.: TRICAAS-1610212022006), and Zhejiang Provincial Natural Science Foundation of China (Grant CRediT authorship contribution statement Hongkai Zhu: Conceptualization, Methodology, Validation, Visualization, Software, Formal analysis, Investigation, acquisition. Li Niu: Formal analysis, Investigation, Methodology. Li Zhu: Formal analysis, Investigation, Methodology. Haibo Yuan: Formal analysis, Investigation, Software, Validation. Paul A. Kilmartin: Formal analysis. Yongwen Jiang: Formal analysis, Investigation, Methodology, Resources, Software, Validation, Visualization.