This work was supported by the National Natural Science Foundation of China (numbers 31672457, 22004035), China Postdoctoral Science Foundation (numbers 2020TQ0094, 2020M682565), Hunan Provincial Science and Technology Department (numbers 2018WK4025, 2020NK2004, 2020ZL2004, 2021JJ40230, 2021JJ30008), the Double first-class construction project of Hunan Agricultural University (numbers SYL201802003, YB2018007, CX20190497), the science and technology innovation Program of Hunan Province (number 2020RC2054).