Conceptualization, S.W. and L.L.; data curation, J.W., J.Z. (Jialei Zhang), Y.G. and J.Z. (Jie Zou); methodology, F.G. and S.Y.; formal analysis, J.W. and J.Z. (Jie Zou); writing—original draft, J.W., Y.G. and J.Z. (Jie Zou); writing—review and editing, J.W. and J.Z. (Jie Zou); funding acquisition, L.L. and S.W. All authors have read and agreed to the published version of the manuscript. This research was funded by the Key R&D Program of Shandong Province, China (2021LZGC026); the Major Science and Technology Innovation Project of Shandong Province, China (2022CXPT031); and the National Modern Agricultural Industrial Technology System Funding Project (CARS-13).