This work was supported by the Natural Science Foundation of Hunan Province (2023JJ30285 and 2020JJ4346), the Foundation of Department of Education of Hunan Province (20K063), the Foundation of Hunan Provincial Department of Agriculture and Rural Affairs ( [2023JJ30285, 2020JJ4346]; Natural Science Foundation of Hunan Province [20K063]; Foundation of Department of Education of Hunan Province; Foundation of Hunan Provincial Department of Agriculture and Rural Affairs (Advantage and Characteristic Industry Cluster and Industry Integration, Early Mature Rapeseed Industry Cluster) [S202210537025X]; Provincial Innovation and Entrepreneurship Training Program for College Students [QL20230166]; Postgraduate Scientific Research Innovation Project of Hunan Province