L.Z. and X.L. (Xin Li) designed the experiment and wrote the original draft; X.L. (Xiangyan Liu), Q.X. and J.Q. analyzed the data and prepared for graph; X.L. (Xiaowen Li), Y.Z., and X.W. prepared the experiment and made all H&E and oil red staining sections; J.W. and L.W. wrote the original draft and reviewed the manuscript; J.W. reviewed the manuscript and checked the grammar; L.W. reviewed the manuscript and provided funding acquisition. All authors have read and agreed to the published version of the manuscript. This research was funded by the Special Funds for Construction of Innovative Provinces in Hunan Province, grant number 2020NK2032 and the Frontier Key Science and Technology Research in the Pig Industry, grant number 2019xny-js044.