S.Y.: Data curation, Formal analysis, Methodology, Writing—original draft. C.C.: Data curation, Formal analysis. Y.H.: Data curation, Formal analysis. P.C.: Methodology, Writing—review and editing. N.T.: Funding acquisition, Project administration, Resources. Z.Y.: Funding acquisition, Supervision, Project administration, Writing—review and editing. All authors have read and agreed to the published version of the manuscript. This work was financially supported by the National Key R&D Program of China “Scientific and Technological Innovation for Increased Grain Yield and Efficiency” Key Project (2017YFD0301501, 2018YFD0301005), and the Hunan Youth Science and Technology Innovation Talent Program (2020RC3073).