Conceptualization, X.L. and X.Z.; Data curation, L.H.; Formal analysis, Y.Z.; Funding acquisition, X.L.; Investigation, Z.Z. and K.L.; Project administration, L.B.; Supervision, X.Z.; Validation, X.Z.; Writing—original draft, X.L.; Writing—review & editing, L.P. and X.Z. This research was funded by the National Natural Science Foundation of China (grant number 31672043 and 21003042), the Natural Science Foundation of Hunan Province (grant number 2018JJ2165), China Postdoctoral Science Foundation funded project (grant number 2015M582329), and China Scholarship Council (grant number 201608430210).