Funding information National Natural Science Foundation of China, Grant/Award Number: 31901905; Natural Science Foundation of Hunan Province, China, Grant/Award Number: 2020JJ5238; Scientific Research Fund of Hunan Provincial Education Department, Grant/Award Number: 19B254; China Agriculture Research System, Grant/Award Number: CARS-16-E19; Central Government Directs Local Science and Technology Development Projects, Grant/Award Number: 2019CT5009; Youth Fund Project of Hunan Agricultural University, Grant/Award Number: 19QN29
Junzhi Wang and Yajun Peng contributed equally to this study.