We thank Dr. Xiao-Wen Cheng (Department of Microbiology, Miami University, United States) for providing the ascovirus materials. Funding. This work was supported by the National Natural Science Foundation of China (31801799), the Hunan Province Natural Science Foundation (2019JJ50274), the Hunan Education Bureau Research Foundation (18B123), the National Natural Science Foundation of China (31872027), Changsha Science and Technology Project (kq1901033), and the Double First-Class Construction Project of Hunan Agricultural University.