Conceptualization, J.Y. and J.W.; methodology, C.L. and C.H.; software, L.Z. and L.K.; validation, C.L., C.H. and L.Z.; formal analysis, Z.Y. and L.W.; investigation, J.W., R.L. and J.Y.; resources, J.Y. and J.W.; data curation, J.Y. and C.L.; writing—original draft preparation, C.L., C.H. and L.Z.; writing—review and editing, J.Y. and L.Z.; visualization, J.W.; supervision, J.Y. and J.W.; project administration, J.Y. and J.W.; funding acquisition, J.Y. All authors have read and agreed to the published version of the manuscript. The research was funded by the Natural Science Foundation of Hunan Province, China (Grant No. 2020JJ4368), “Double First-Class” Disciplinary Construction Project of Hunan Agricultural University (SYL2019049) and Innovation Research and Development Project of the Hunan Development and Reform Commission (37).