This study was financially supported by the Hunan Province Science Fund for Excellent Young Scholars (2021JJ20030), the National Natural Science Foundations of China (42177026), the Central Financial Project (JXTG(2020)25), the Forestry Science and Technology Innovation (2023(10)), the Key Research and Development Programs of Hunan (2020NK2011 and 2021NK2012), and the Double First-class Construction Project of Hunan Agricultural University (kxk201801007 and SYL2019025).