Conceptualization, Q.Z., H.G. and Y.Y.; methodology, H.G. and K.X.; software, Q.Z., G.S. and X.D.; validation, H.G. and S.Y.; formal analysis, Q.Z. and F.Y.; investigation, Y.P., Y.C. and X.H.; resources, J.H., Y.Y. and K.X.; data curation, F.Y., Y.F., Q.W., S.Y. and Z.J.; writing—original draft preparation, Q.Z., H.G., S.Y. and F.Y.; writing—review and editing, J.H., Y.P., Y.Y. and K.X. visualization, H.G. and S.Y.; supervision, Y.Y. and K.X.; project administration, Y.Y. and K.X.; funding acquisition, Y.Y. and K.X. All authors have read and agreed to the published version of the manuscript. This research was funded by the Laboratory of Lingnan Modern Agriculture Project (NT2021005), the Strategic Priority Research Program of Chinese Academy of Sciences (Grant No. XDA24030204), the Special Funds for the Construction of Innovative Provinces in Hunan (Grant numbers 2021NK1009 and 2021NK1012); the Natural Science Foundation of Hunan Province Project (2023JJ20043 and 2020JJ5635).