Conceptualization, H.K. and Y.X. (Yuchen Xu); writing—original draft, Y.X. (Yuchen Xu); writing—review & editing, H.K. and Y.X. (Yuchen Xu); funding acquisition, H.K. and S.Y.; formal analysis, H.K., Y.X. (Yuchen Xu), Y.L., S.P. and Q.L.; investigation, Y.X. (Yuchen Xu), S.Y., S.J., L.B. and R.C.; visualization, Y.X. (Yuchen Xu) and L.B.; validation, Y.X. (Yuchen Xu) and L.B.; project administration, H.K. and Y.X. (Yinghui Xiao). All authors have read and agreed to the published version of the manuscript. This work was financially supported by the International Science and Technology Innovation Program of the Chinese Academy of Agricultural Sciences (CAASTIP) (CAAS-ZDRW202108), the National Natural Science Foundation of China to Houxiang Kang (32261143468 & 31772120), and the Key Technologies R&D Program of Tianjin to Shuangyong Yan (18YFZCNC01210).