Conceptualization, X.Z. and J.Y.; methodology, X.Z.; software, F.H.; validation, Y.G., F.H. and Z.D.; formal analysis, Z.D.; investigation, F.H.; resources, Z.X.; data curation, Y.G.; writing—original draft preparation, Y.G.; writing—review and editing, X.Z.; visualization, Y.G.; supervision, J.Y. and X.Z.; project administration, X.Z.; funding acquisition, G.G. All authors have read and agreed to the published version of the manuscript. This work was supported by the Hunan Provincial Science and Technology Department (2021JJ30320), Hunan Provincial Department of Education (23A0186), the Science and Technology Innovation Program of Hunan Province (2023RC1074), and the Youth Innovation Promotion Association CAS and China Agriculture Research System of MOF and MARA (CARS-35).
