Conceptualization, J.L. and X.Z.; investigation, Z.L. and H.M.; resources, C.Z., Y.M. and A.Z.; data curation, H.M. and Y.Y.; writing—original draft preparation, J.L., X.C. and H.M.; writing—review and editing, J.L. and H.M.; supervision, H.Z., W.Z. and Y.Z.; funding acquisition, J.L., H.M. and X.Z. All authors have read and agreed to the published version of the manuscript. This research was funded by the National Natural Science Foundation of China (31701815), the Natural Science Foundation of Hunan (2021JJ40288), the Changsha Municipal Natural Science Foundation (kq2007018), and Hunan Agricultural Science and Technology Innovation Fund Project (2021CX85 and 2021CX87).