The authors thank Mr. Yu-Heng Wu, Mr. Ming-Wei Liao, Mr. Bin Chen and Miss Min Deng (Hunan Agricultural University, China), Mr. Wen-Wu Zhang, Mr. Min Wang, and Mr. Ming-Jian Wang (Hunan Wuyunjie National Nature Reserve Administration), Mr. Gui-Qing Yu and Miss Xin Li (Hunan Hupingshan National Nature Reserve Administration), Mr. Chun-Lin Liao (Hunan Badagongshan National Nature Reserve Administration) and Mr. Bi-Wu Qin (Tianpingshan, Badagongshan National Nature Reserve, Hunan) for their help in the field. This study was supported by the Biodiversity Survey and Assessment Project of the Ministry of Ecology and Environment, China ( 2019HJ2096001006 ) and Double first-class construction project of Hunan Agricultural University .