Conceptualization, D.W., N.W.; methodology, D.W., J.M.; software, D.W., J.M.; validation, D.W., J.M.; formal analysis, D.W., J.M.; investigation, D.W.; resources, D.W.; data curation, D.W.; writing—original draft preparation, D.W., N.W.; writing—review and editing, N.W., Y.Y.; visualization, D.W.; supervision, N.W., Y.Y.; project administration, N.W., Y.Y.; funding acquisition, D.W., N.W. All authors have read and agreed to the published version of the manuscript. This research was funded by Hunan Provincial Natural Science Foundation of China, grant number 2018JJ2177; Double first-class construction project of Hunan Agricultural University, grant number SYL2019048 and Postgraduate Research and Innovation Project of Hunan Province, grant number CX2018B394.