We thank the Hunan Province Science Foundation for Youths (2018JJ3215), Research Foundation of Education Bureau of Hunan Province (18C0147) and Hunan Agricultural University Science Foundation for High-Level Personnel (17QN21), Double first-class construction project of Hunan Agricultural University. We are grateful for the financial support from the National Natural Science Foundation of China (No. 31672457), Local Science and Technology Development Project guided by the Central Government (YDZX20184300002303), Ministry of Agriculture of the People’s Republic of China (2015-Z64 and 2016-X47) and Hunan Provincial Science and Technology Department (2016NK2101, 2016WK2008, 2016TP2005, and 2018CT5002).